Acids are produced from L-arabinose, D-glucose, m-inositol and sucrose, but not from D
-mannitol, L-rhamnose or D-sorbitol. The predominant ubiquinone is Q-9. The major fatty acids are C18:1x7c, C16:0, C16:1x7c and C19:0cyclo x8c. The DNAG?C content of the type strain is 61.3 ± 2.1 %